Captured by smugglers when he was just a hatchling, a macaw named Blu never learned to fly and lives a happily domesticated life in Minnesota with his human friend, Linda. Blu is thought to be the last of his kind, but when word comes that Jewel, a lone female, lives in Rio de Janeiro, Blu and Linda go to meet her. Animal smugglers kidnap Blu and Jewel, but the pair soon escape and begin a perilous adventure back to freedom -- and Linda.
When Sid accidentally destroys Manny's heirloom Christmas rock and ends up on Santa's naughty list, he leads a hilarious quest to the North Pole to make things right and ends up making things much worse. Now it's up to Manny and his prehistoric posse to band together and save Christmas for the entire world!
Times are changing for Manny the moody mammoth, Sid the motor mouthed sloth and Diego the crafty saber-toothed tiger. Life heats up for our heroes when they meet some new and none-too-friendly neighbors – the mighty dinosaurs.